phrog_1870 is made of 81 protein sequences and is annotated as "exonuclease"
Functional category: DNA, RNA and nucleotide metabolism


Average length of proteins Number of columns in the MSA Number of conserved columns in the MSA
311.89 353 315


List of all proteins of this PHROG

PHROG prot ID NCBI prot ID NCBI prot annotation Virus ID (click to view) Virus name Virus taxonomy
NC_022336_p36 YP_008531671.1 putative exonuclease NC_022336 Propionibacterium phage PHL010M04 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_027295_p36 YP_009145850.1 exonuclease NC_027295 Propionibacterium phage PHL199M00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027389_p36 YP_009152626.1 exonuclease NC_027389 Propionibacterium phage PHL141N00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027624_p36 YP_009160043.1 exonuclease NC_027624 Propionibacterium phage SKKY Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_022337_p36 YP_008531716.1 putative exonuclease NC_022337 Propionibacterium phage PHL071N05 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_022341_p36 YP_008531898.1 putative exonuclease NC_022341 Propionibacterium phage PHL113M01 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027373_p36 YP_009151554.1 exonuclease NC_027373 Propionibacterium phage PHL030N00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
MG820641_p35 AUV62445.1 exonuclease MG820641 Propionibacterium phage pa615 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
MF919515_p36 ATN90315.1 exonuclease MF919515 Propionibacterium phage Leviosa Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
MG820632_p36 AUV62068.1 exonuclease MG820632 Propionibacterium phage pa310 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_022340_p36 YP_008531852.1 putative exonuclease NC_022340 Propionibacterium phage PHL114L00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_009541_p37 YP_001285613.1 gp37 NC_009541 Propionibacterium phage PA6 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_031005_p36 YP_009278014.1 exonuclease NC_031005 Propionibacterium phage QueenBey Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_027385_p36 YP_009152397.1 exonuclease NC_027385 Propionibacterium phage PHL092M00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
MF919505_p36 ATN89087.1 exonuclease MF919505 Propionibacterium phage DrParker Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_027386_p36 YP_009152442.1 exonuclease NC_027386 Propionibacterium phage PHL152M00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027623_p36 YP_009159999.1 exonuclease NC_027623 Propionibacterium phage Pirate Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_031119_p36 YP_009292071.1 exonuclease NC_031119 Propionibacterium phage Enoki Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
MF919491_p36 ATN87133.1 exonuclease MF919491 Propionibacterium phage Aquarius Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
MF919516_p36 ATN90359.1 exonuclease MF919516 Propionibacterium phage LilBandit Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_027367_p35 YP_009151235.1 exonuclease NC_027367 Propionibacterium phage PHL132N00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_028967_p35 YP_009214910.1 putative exonuclease NC_028967 Propionibacterium phage PAC1 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_027359_p36 YP_009150041.1 exonuclease NC_027359 Propionibacterium phage PHL082M00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
MG820637_p35 AUV62278.1 exonuclease MG820637 Propionibacterium phage pa63 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_018841_p36 YP_006906576.1 hypothetical protein NC_018841 Propionibacterium phage P101A Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027361_p36 YP_009150235.1 exonuclease NC_027361 Propionibacterium phage PHL085N00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
MG820635_p36 AUV62193.1 exonuclease MG820635 Propionibacterium phage pa29399-1-D_1 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_027401_p36 YP_009153285.1 exonuclease NC_027401 Propionibacterium phage PHL095N00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
JX570706_p36 AGI12577.1 putative exonuclease JX570706 Propionibacterium phage PHL037M02 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; Propionibacterium virus PHL037M02.
MG820636_p36 AUV62236.1 exonuclease MG820636 Propionibacterium phage pa29399-1-D_2 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_018852_p36 YP_006907165.1 hypothetical protein NC_018852 Propionibacterium phage P100D Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
MF919522_p36 ATN90931.1 exonuclease MF919522 Propionibacterium phage MEAK Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_031003_p36 YP_009277910.1 exonuclease NC_031003 Propionibacterium phage Moyashi Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_027391_p36 YP_009152697.1 exonuclease NC_027391 Propionibacterium phage PHL041M10 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_015453_p36 YP_004414741.1 Gp37 NC_015453 Propionibacterium phage PAS50 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
JX570703_p36 AGI12441.1 putative exonuclease JX570703 Propionibacterium phage PHL073M02 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_018851_p36 YP_006907119.1 hypothetical protein NC_018851 Propionibacterium phage ATCC29399B_C Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
MG820643_p37 AUV62533.1 exonuclease MG820643 Propionibacterium phage pa35 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_027620_p36 YP_009159863.1 exonuclease NC_027620 Propionibacterium phage MrAK Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
MG820640_p35 AUV62404.1 exonuclease MG820640 Propionibacterium phage pa15 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_028694_p34 YP_009191379.1 putative exonuclease NC_028694 Propionibacterium phage PA1-14 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_022342_p36 YP_008531942.1 putative exonuclease NC_022342 Propionibacterium phage PHL111M01 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_018834_p36 YP_006906062.1 RecB NC_018834 Propionibacterium phage P9.1 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
MG820642_p35 AUV62486.1 exonuclease MG820642 Propionibacterium phage pa28 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_027336_p36 YP_009147260.1 exonuclease NC_027336 Propionibacterium phage PHL009M11 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_018838_p36 YP_006906437.1 hypothetical protein NC_018838 Propionibacterium phage P100_A Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
KY926792_p8 ASJ79904.1 putative exonuclease KY926792 Propionibacterium phage PacnesP1 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_027371_p11 YP_009151434.1 hypothetical protein NC_027371 Propionibacterium phage Pacnes 2012-15 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_022335_p36 YP_008531626.1 putative exonuclease NC_022335 Propionibacterium phage PHL067M10 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_018849_p36 YP_006907033.1 hypothetical protein NC_018849 Propionibacterium phage P105 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027625_p36 YP_009160090.1 exonuclease NC_027625 Propionibacterium phage Kubed Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
MG820638_p35 AUV62321.1 exonuclease MG820638 Propionibacterium phage pa6919-4 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_027630_p36 YP_009160314.1 exonuclease NC_027630 Propionibacterium phage Ouroboros Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_018845_p36 YP_006906808.1 hypothetical protein NC_018845 Propionibacterium phage P104A Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027294_p36 YP_009145805.1 exonuclease NC_027294 Propionibacterium phage PHL150M00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_018839_p36 YP_006906482.1 hypothetical protein NC_018839 Propionibacterium phage P14.4 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_031084_p36 YP_009288358.1 exonuclease NC_031084 Propionibacterium phage BruceLethal Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
MF919533_p36 ATN91976.1 exonuclease MF919533 Propionibacterium phage Supernova Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_027370_p36 YP_009151414.1 exonuclease NC_027370 Propionibacterium phage PHL179M00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027346_p36 YP_009148307.1 exonuclease NC_027346 Propionibacterium phage PHL171M01 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027354_p36 YP_009149552.1 exonuclease NC_027354 Propionibacterium phage PHL301M00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027357_p36 YP_009149950.1 exonuclease NC_027357 Propionibacterium phage PHL025M00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027333_p36 YP_009146929.1 exonuclease NC_027333 Propionibacterium phage PHL070N00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027628_p36 YP_009160224.1 exonuclease NC_027628 Propionibacterium phage Lauchelly Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_022334_p36 YP_008531580.1 putative exonuclease NC_022334 Propionibacterium phage PHL112N00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_018842_p36 YP_006906623.1 hypothetical protein NC_018842 Propionibacterium phage P1.1 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027400_p36 YP_009153240.1 exonuclease NC_027400 Propionibacterium phage PHL055N00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_027627_p36 YP_009160179.1 exonuclease NC_027627 Propionibacterium phage Solid Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
MG820634_p35 AUV62150.1 exonuclease MG820634 Propionibacterium phage pa27 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_027622_p36 YP_009159953.1 exonuclease NC_027622 Propionibacterium phage Stormborn Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027629_p36 YP_009160269.1 exonuclease NC_027629 Propionibacterium phage Attacne Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027362_p36 YP_009150279.1 exonuclease NC_027362 Propionibacterium phage PHL116M00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_027621_p36 YP_009159909.1 exonuclease NC_027621 Propionibacterium phage Wizzo Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
KR337649_p36 AKO60490.1 exonuclease KR337649 Propionibacterium phage Keiki Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; Propionibacterium virus Keiki.
NC_022338_p36 YP_008531761.1 putative exonuclease NC_022338 Propionibacterium phage PHL060L00 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
MG820645_p36 AUV62617.1 exonuclease MG820645 Propionibacterium phage pa3-SS3 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus; unclassified Pahexavirus.
NC_018847_p36 YP_006906932.1 hypothetical protein NC_018847 Propionibacterium phage ATCC29399B_T Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_015454_p36 YP_004414787.1 Gp37 NC_015454 Propionibacterium phage PAD20 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
NC_018840_p36 YP_006906529.1 hypothetical protein NC_018840 Propionibacterium phage P100_1 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Siphoviridae; Pahexavirus.
KU886273_p4 ANH49483.1 PD-(D/E)XK nuclease superfamily protein KU886273 Freshwater phage uvFW-CGR-AMD-COM-C455 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Podoviridae; unclassified Podoviridae.
KU886269_p55 ANH49319.1 PD-(D/E)XK nuclease superfamily protein KU886269 Freshwater phage uvFW-CGR-AMD-COM-C403 Viruses; Duplodnaviria; Heunggongvirae; Uroviricota; Caudoviricetes; Caudovirales; Podoviridae; unclassified Podoviridae.

PHROGs colocalized to this PHROG

PHROG #Prot Annotation Category #Neighbors Probability Score
phrog_801 180 endolysin; inhibits RNA polymerase lysis 79 3.54E-258 248.57
phrog_1890 80 no annotation unknown function 79 1.19E-310 301.05

PHROGs similar to this PHROG

PHROG Annotation Category Probability Evalue posQ/posS
phrog_1218 exonuclease DNA, RNA and nucleotide metabolism 98.1 4.5E-09 16-222 / 10-205
phrog_1706 CRISPR/Cas system associated DNA, RNA and nucleotide metabolism 96.8 8.3E-06 158-289 / 90-225
phrog_298 exonuclease DNA, RNA and nucleotide metabolism 94.9 0.0016 164-219 / 126-187
phrog_412 exonuclease VIII DNA, RNA and nucleotide metabolism 93.3 0.01 16-263 / 22-245
phrog_466 exonuclease DNA, RNA and nucleotide metabolism 95.9 0.0002 152-251 / 240-353
phrog_69 exonuclease DNA, RNA and nucleotide metabolism 99.4 4.4E-17 14-293 / 7-248

PFAMs similar to this PHROG

ID Definition Evalue posQ/posS
PF12705.6 PD-(D/E)XK nuclease superfamily 5.6E-21 172-293 / 129-243
PF06023.11 CRISPR-associated exonuclease Csa1 2.2E-19 21-293 / 46-281
PF10926.7 Protein of unknown function (DUF2800) 1.1E-18 22-297 / 3-231
PF12684.6 PDDEXK-like domain of unknown function (DUF3799) 3.9E-17 24-70 / 1-52
PF09810.8 Exonuclease V - a 5' deoxyribonuclease 5.9E-17 21-69 / 12-60
PF01930.16 Domain of unknown function DUF83 2.5E-16 176-293 / 53-159
PF08696.10 DNA replication factor Dna2 3.3E-10 15-70 / 58-115

Annotations of the NCBI proteins of this PHROG


66:exonuclease 11:hypothetical protein 2:PD-(D/E)XK nuclease superfamily protein 1:RecB 1:similar to Bxb1 gp62

KEGG orthologous groups similar to this PHROG

ID Name : Definition Pathways BestScore #Hits
K07465 K07465 : putative RecB family exonuclease no pathways 76 6

GO terms in Uniprot proteins similar to this PHROG

ID Definition
GO:0045145 single-stranded DNA 5'-3' exodeoxyribonuclease activity

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